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A number of factors that address both ecological conditions (such as forests, soils, climate, and hydrology) and human impacts (distance from roads, agriculture, forest loss, and density of “forest edge”) were tested, producing a spatial model that identified and mapped the most important environmental factors contributing to bonobo occurrence. The most important predictor of bonobo presence is distance from agricultural areas, according to the researchers. They also discovered that only 27.5 percent of the suitable habitat is located in areas that are currently protected.